Corrected entry: Doctors can't just randomly work in any department they feel like. Cameron, an immunologist, is not qualified to work in the ER. Chase, on the other hand, is, as he is specialised in emergency medicine, but he works on the surgical staff despite being neither a surgeon nor an anesthesiologist.
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Correction: It's stated that Chase works as some kind of assistant in the surgical team, the surgeons are referred to as his bosses. With Chase having his time in House's team on his resumée (House obviously has quite a reputation) he would have no problem getting a job like that. As for Cameron working in the ER, well, not many doctors want to work in busy ERs so they pretty much take what they can get, even doctors that technically aren't qualified (or so my mother tells me, she works in one). Also, again, working with a famous doctor like House gives advantages.
littlestar